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Order, to each of the two dna strands entwined in the double helix. as the single fertilized egg cell grows into an adult, the genome is replicated many billions of times, its precise sequence of three billion nucleotides retained with extraordinary fidelity. the rate of error -that is, the insertion into the newly made dna sequence of a nucleotide out of its proper order -- is about one in 10 billion nucleotides. but on its own, dna is incapable of such faithful replication; in a test-tube experiment, a dna strand, provided with a mixture of its four constituent nucleotides, will line them up with about one in a hundred of them out its proper place. on the other hand, when the appropriate protein enzymes are added to the test tube, the fidelity with which nucleotides are incorporated in the newly made dna strand is greatly improved, reducing the error rate to one in 10 million. these remaining errors are finally reduced to one in 10 billion by a set of "repair" enzymes also proteins ; that detect and remove mismatched nucleotides from the newly synthesized dna. thus, in the living cell the gene's nucleotide code can by replicated faithfully only because an array of specialized proteins intervenes to prevent most of the errors -- which dna by itself is prone to make -- and to repair the few remaining ones. moreover, it has been known since the 1960s that the enzymes that synthesize dna influence its nucleotide sequence. in this sense, genetic information arises not from dna alone but through its essential collaboration with protein enzymes -- a contradiction of the central dogma's precept that inheritance is uniquely governed by the self-replication of the dna double helix. another important divergent observation is the following: in order to become biochemically active and actually generate the inherited trait, the newly made protein, a strung-out ribbon of a molecule, must be folded up into a precisely organized ball-like structure. the biochemical events that give rise to genetic traits -- for example, enzyme action that synthesizes a particular eye-color pigment -- take place at specific locations on the outer surface of the three-dimensional protein, which is created by the particular way in which the molecule is folded into that structure. to preserve the simplicity of the central dogma, crick was required to assume, without any supporting evidence, that the nascent protein -- a linear molecule -- always folded itself up in the right way once its amino acid sequence had been determined. in the 1980s, however, it was discovered that some nascent proteins are on their own likely to become misfolded -- and therefore remain biochemically inactive -- unless they come in contract with a special type of "chaperone" protein that properly folds them. the importance of these chaperones has been underlined in recent years by research on degenerative brain diseases that are caused by "prions, " research that has produced some of the most disturbing evidence that the central dogma is dangerously misconceived. crick's theory holds that biological replication, which is essential to an organism's ability to infect another organism, cannot occur without nucleic acid. yet when scrapie, the earliest known such disease, was analyzed biochemically, no nucleic acid -- neither dna nor rna -- could by found in the infectious material that transmitted the disease. in the 1980's, stanley prusiner confirmed that the infectious agents that cause scrapie, mad cow disease, and similar very rare but invariably fatal human diseases are indeed nucleic-acid-free proteins he named them prions ; , which replicate in an entirely unprecedented way. invading the brain, the prion encounters a normal brain protein which it then refolds to match the prion's distinctive three-dimensional shape. the newly refolded protein itself becomes infectious and, acting on another molecule of the normal protein, sets up a chain reaction that propagates the disease to its fatal end.
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Rsum la fin des annes 1890, les mutoscopes taient associs aux productions dites pour adultes. pourtant, ces appareils taient souvent accessibles aux enfants. deux sources de documentation, la campagne du groupe de presse hearst, en 1899, contre les images des galeries de l'enfer , ainsi que les photographies de la farm security administration, remontant aux annes 1930, permettent d'tablir la longvit de la frquentation des mutoscopes par la jeunesse amricaine. la panique morale qu'instillait la rhtorique des journaux de hearst fut de courte dure et servit surtout les intrts du groupe de presse. elle demeure cependant fort instructive pour qui s'intresse la manire dont les discours sur les nouveaux mdias diffusent une mme angoisse au cours de tout le vingtime sicle. au contraire, les photographies du fsa rvlent pour leur part que les indcences du mutoscope n'taient ressenties comme telles que de faon marginale dans les annes 1930. abstract by the late 1890s, mutoscopes were associated with "adult" material. yet such displays were often accessible to children. two sources of documentation, an 1899 hearst newspaper campaign against "picture galleries of hell" and u.s. farm security administration photographs of the 1930s, demonstrate the longevity of american children's access to mutoscopes. the rhetoric of moral panic used in the hearst campaign was shortlived and self-interested, but instructive in demonstrating how discourse on new media displays a similarly anxious tenor throughout the twentieth century. by contrast, the fsa photographs reveal that the perceived and play texas holdem poker online.

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